... =a12J(1 ,2) + J (2, 2) = r2Q2 + a2QQ2 J (2, 2) 76 dS1 = F = J (1,1) +a21J (2, 1) +a31J (3,1) - J (1 ,2) dt dS2 = F2 = J (2, 2) +a12J (1 ,2) - J (2, 1) - J (2, 3) dt dS = F3 = J (3,3) + a 23 J ( 2, 3) - J (3,1) ... K - S1 J (1,1) = r ç ÷ S1 è K ø 66 Mồi S1 Q1,N J(1 ,2) Thú S2 Q2,P J (2, 2) J(1,1) Q1 Q2 J (1,1) º r1Q1 J (1 ,2) º aQ1Q2 J (2, 2) º r2Q2 67 J(3,1) Dinh dưỡng S1 J(1,1) J(1 ,2) J (2, 1) Thực vật S2 J (2, 2) ... dS1 S1 a2 K2 p ÷S2 S3 ú + = F1 = qD1 (v) - q + a 21 S2GT ( X T ) + a31 ê K3S3 + a 23 ç a ç dt V K + S2 ÷ è ø ë û a1K1S1S2 GT ( X T )GS ( X S )M K1 + S1 dS2 S2 a1K1S1S = F2 = qC2 - q + a 12 MGT (...